Mammoth
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Order: Proboscidea
Family: Elephantidae
Genus: Mammuthus

[Photo] A model of a mammoth in Nordhausen, Gemany. pic made by J C D (http://commons.wikimedia.org/wiki/User:J_C_D). Public domain by the author. Photo taken on 27 March 2006.

A mammoth is any species of the extinct genus Mammuthus. These proboscideans (elephants or their extinct relatives) were often equipped with long curved tusks and, in northern species, a covering of long hair. They lived from the Pliocene epoch from 4.8 million years ago to around 4,500 years ago. The word mammoth comes from the Russian мамонт mamont, probably in turn from the Vogul (Mansi) language.

Species
- Mammuthus africanavus, African mammoth
- Mammuthus columbi, Columbian mammoth
- Mammuthus exilis, Pygmy mammoth
- Mammuthus imperator, Imperial mammoth
- Mammuthus jeffersonii, Jeffersonian mammoth
- Mammuthus trogontherii, Steppe mammoth
- Mammuthus meridionalis, Southern mammoth
- Mammuthus subplanifrons, South African mammoth
- Mammuthus primigenius, Woolly mammoth
- Mammuthus lamarmorae, Sardinian dwarf mammoth
- Mammuthus sungari, Songhua River Mammoth

Evolutionary history
Mammoth remains have been found in Europe, Africa, Asia, and North America. They are believed to have originally evolved in North Africa about 4.8 million years ago, during the Pliocene, where bones of Mammuthus africanavus have been found in Chad, Libya, Morocco and Tunisia. Mammuthus subplanifrons, found in South Africa and Kenya, is also believed to be one of the oldest species (about 4 million years ago).

Despite their African ancestry, they are in fact more closely related to the modern Asian Elephant than either of the two African elephants (as both Mammuthus and Elephas also originated in Africa). The common ancestor of both mammoths and Asian elephants split from the line of African elephants in the Late Miocene about 6 - 7.3 million years ago, probably due to the uplift of East Africa and increasing aridity in the Middle East. The Asian elephants and mammoths diverged about half a million years later, i.e. 5.5 - 6.3 million years ago.(Capelli et al. 2006)

In due course the African mammoth migrated north to Europe and gave rise to a new species, the southern mammoth (Mammuthus meridionalis). This eventually spread across Europe and Asia and crossed the now-submerged Bering Land Bridge into North America.

Around 700,000 years ago, the warm climate of the time deteriorated markedly and the savannah plains of Europe, Asia and North America gave way to colder and less fertile steppes. The southern mammoth consequently declined, being replaced across most of its territory by the cold-adapted steppe mammoth (Mammuthus trogontherii). This in turn gave rise to the woolly mammoth, Mammuthus primigenius) around 300,000 years ago. Woolly mammoths were better able to cope with the extreme cold of the Ice Ages.

The woolly mammoths were a spectacularly successful species; they ranged from Spain to North America and are thought to have existed in massive numbers. The Russian researcher Sergei Zimov estimates that during the last Ice Age, parts of Siberia may have had an average population density of sixty animals per hundred square kilometres - equivalent to African elephants today.

Extinction
Most mammoths died out at the end of the last Ice Age. A definitive explanation for their mass extinction is yet to be agreed upon. A small population survived on St. Paul Island, Alaska, up until 6000 BC, and the small mammoths of Wrangel Island became extinct only around 2000 BC

Whether the general mammoth population died out for climatic reasons or due to overhunting by humans is controversial. Another theory suggests that mammoths may have fallen victim to an infectious disease. A combination of climate change and hunting by humans is probably the most likely explanation for their extinction.

New data derived from studies done on living elephants (see Levy 2006) suggests that though human hunting may not have been the primary cause toward the mammoth's final extinction, human hunting was likely a strong contributing factor. Homo erectus is known to have consumed mammoth meat as early as 1.8 million years ago (Levy 2006: 295).

However, the American Institute of Biological Sciences also notes that bones of dead elephants, left on the ground and subsequently trampled by other elephants, tend to bear marks resembling butchery marks, which have previously been misinterpreted as such by archaeologists.

The survival of the dwarf mammoths on Russia's Wrangel Island was due to the fact that the island was very remote, and uninhabited in the early Holocene period. The actual island was not discovered by modern civilization until the 1820s by American whalers. A similar dwarfing occurred with the Pygmy Mammoth on the outer Channel Islands of California, but at an earlier period. Those animals were very likely killed by early Paleo-Native Americans, and habitat loss caused by a rising sea level that split the Santa Rosae into the outer Channel Islands.

Size
It is a common misconception that mammoths were much larger than modern elephants, an error that has led to "mammoth" being used as an adjective meaning "very big". Certainly, the largest known species, the Imperial Mammoth of California, reached heights of at least 5 metres (16 feet) at the shoulder. Mammoths would probably normally weigh in the region of 6 to 8 tonnes, but exceptionally large males may have exceeded 12 tonnes. A 3.3 m. (11 ft.) long mammoth tusk was discovered north of Lincoln, Illinois in 2005. However, most species of mammoth were only about as large as a modern Asian Elephant. Fossils of species of dwarf mammoth have been found on the Californian Channel Islands (Mammuthus exilis) and the Mediterranean island of Sardinia (Mammuthus lamarmorae). There was also a race of dwarf woolly mammoths on Wrangel Island, north of Siberia, within the Arctic Circle.

Based on studies of their close relatives, the modern elephants, mammoths probably had a gestation period of 22 months, resulting in a single calf being born. Their social structure was probably the same as that of African and Asian elephants, with females living in herds headed by a matriarch, whilst bulls lived solitary lives or formed loose groups after sexual maturity.

Well-preserved specimens
In May of 2007, the carcass of a six-month-old female mammoth calf was discovered encased in a layer of permafrost near the Yuribei River in Russia where it had been buried for 37,000 years. Alexei Tikhonov, the Russian Academy of Science's Zoological Institute's deputy director has dismissed the prospect of cloning the animal, as the whole cells required for cloning would have burst under the freezing conditions. DNA is expected to be well-preserved enough to be useful for research on mammoth phylogeny and perhaps physiology however.

http://en.wikipedia.org/wiki/Mammoth

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