Snow Petrel
From Wikipedia, the free encyclopedia

Order: Procellariiformes
Family: Procellariidae

[Photo] Snow Petrel (Pagodroma nivea). Author: Kristan Hutchison - Antarctic Sun Staff. Source: http://antarcticsun.usap.gov/oldissues2000-2001/2000_1224/petrels.html

The Snow Petrel (Pagodroma nivea) is a small, pure white fulmarine petrel with black underdown, coal-black eyes, small black bill and bluish gray feet. Body length is 36 to 41 centimeters (14???16 in) and the wingspan is 76 to 79 centimeters. In Du Cane Godman, (Monograph of the Petrels, fasc. 4, 1909), it was figured in an illustration by John Gerrard Keulemans (pl. 73).

(30???31 in). Flight is more fluttering than most petrels. It is the only member of the genus Pagodroma. It is one of only three birds that breed exclusively in Antarctica and has been seen at the South Pole. It has the most southernly breeding distribution of any bird.

There are two subspecies, Pagodroma nivea confusa and Pagodroma nivea nivea. They differ in size, and the greater form has a stouter, larger beak. The greater form has a more resricted distribution--the Adelie Land and Balleny Is. area of Antarctica.

Breeding occurs in colonies on the Antarctic Peninsula, the Antarctic continent and various antarctic islands. Nesting is colonial in small to large colonies on cliffs, usually near the sea, but also inland. Some birds remain at the colony all year, but the main influx at colonies is from the mid-September until early November. Nests are simple pebble-lined scrapes usually in a deep rock crevices with overhanging protection. One white egg is laid in late November to mid-December. The egg is incubated for 41 to 49 days and the chick is brooded for 8 days. They fledge 7 weeks later in late February to mid-May. Colonies are also the sites of cleaning areas where Snow Petrels, far from the sea, bathe in snow. Snow Petrels feed mainly on fish, some cephalopods, mollusks and krill as well as carrion. During the winter they disperse to the pack ice, ice floes and the open sea. Flocks are characteristically seen sitting on icebergs. Only very rarely are they observed north of the pack ice.

Like many petrels these birds squirt waxy, yellowish stomach oil at nest intruders. This oil stinks of fish and is extremely difficult to remove. They are known to live 14 to 20 years.

Taxonomy and nomenclature
The issue of taxonomy and nomenclature for this species is complex, and represents an example of many of the unusual problems which arise out of the varied use of the terminiology.

The snow petrel was first named by George Forster, during Captain Cook's voyages in 1777 (A Voyage Round the World), as Procellaria nivea. As Forster's mention of the petrel was only an observation, the authority of the name fell to Gmelin, in his Systema Naturae (1789). Incidentally, a skin was procured by Cook, who noted this in his own account (1777). It was J. R. Forster who painted a picture of it, and made measurements as well, but only the latter were published, posthumously, by M. H. K. Lichtenstein in 1844. While Gmelin was recognized as the authority for Pagodroma nivea until the early twentieth century, the evidence has allowed for Forster to be construed as such, as he had created the name. In addition, Latham (1785) had described the type, which was in the Leverian Museum, but did not invoke Forster's scientific name for it.

Charles Lucien Bonaparte, in his Conspectus Generum Avium, volume II (1857), designated two subspecific binomial terms with no further descriptions to them: Pagodroma major and Pagodroma minor. Contrary to Mathews in Novitates Zoologicae (1934), the two names do not represent alternative, new names for Pagodroma nivea, but are subspecies, as well as nomen nudum, on account of their being listed similar to how other subspecies Bonaparte listed in the Conspectus. It was also Bonaparte who created the generic name Pagodroma, in the Paris academic journal Compte Rendus.. (1856).

Hermann Schlegel, in his treatise on birds in the Leiden Museum in Mus. Pay-Bas (1863), designated two forms of the snow petrel for the first time. He had believed, erroneously, that the Cook expedition specimen represented the greater snow petrel, and designated it as Pagodroma nivea, incorporating Pagodroma major (Bonaparte) as a synonym of it. He thus redundantly described Pagodroma minor, also from Bonaparte's nomen nudum, as the lesser form. With the measurements provided by Lichtenstein taken from J. R. Forsters' manuscript, it was already clear by this time that nivea represented the lesser snow petrel.

This matter was ultimately clarified by Gregory Mathews, who in Birds of Australia, volume 2 (1912), described the greater form as Pagodroma confusa. From that time debate has persisted as to whether the snow petrel represents one monotypic species, one species with two subspecies, or two different species. In addition, J. Prevost (Oiseaux, 1969) ascribed the name Pagodroma nivea major (Bonaparte) to a specimen of the greater snow petrel.

In the 1850s the "Rattlesnake" expedition procured specimens of the snow petrel from the South Georgia/Falklands Is. region, near South America. A German Antarctic exploring expedition, 1882-83, also obtained skins, and opinion had emerged that the petrel populations there were unique from Pagodroma nivea. What distinguished the skins were the black shafts to the remiges, or flight feathers. Heinrich Pagenstecher in 1885 proposed the binomial name for a subspecies, Pagodroma novegeorgica (Jahrbuch Wiss. Hamburg II). The publication of this name was overlooked by English-speaking naturalists, and a similar account by Karl von der Steinen (1890) which had informally dropped the name novegeorgica, was not; this later account came to represent the scientific authority for South Georgia petrels. Gregory Mathews, in his Birds, designated the type from the "Rattlesnake" expedition to Steinen's "name". Later adding to the confusion, he assumed that the name novegeorgica could not be used for South Georgia petrels: Steinen's use of the term was not valid in a taxonomic sense, and this same term had been reinforced by its usage within the synopses of other, previous works. Thus, he designated a name for the type he designated in Birds, and described the form Pagodroma nivea falklandica (Bulletin of the British Orn. Club, 1926). Eventually, Hellmayr and Conover (Museum of Nat. Hist. Publications, Chicago; 1949) unravelled the matter by recognizing Pagenstecher as the authority for novegeorgica, and omitting reference to Steinen.

In the decade following the United States Exploring Expedition, Titian Ramsay Peale (1848) wrote an account of the bird and mammal specimens collected, including the description of an atypical snow petrel that was completely white--even the loral area behind the eye. The name Procellaria candida was published as an alternative synonym for the entry of Procellaria nivea. Mathews (1912) recognized candida as a subspecies of nivea; however, he later amended this with his description Pagodroma nivea pealei (Bulletin of the British Orn. Club, 1928), as Peale's name did not constitute a valid one. Mathews also described the subspecies Pagodroma nivea alba (Bulletin of the British Orn. Club, 1928) for the type he designated in his book, Birds of Norfolk and Lord Howe Islands (1928); the latter had also included information on the avifauna of the Australasian quadrant of Antarctica.

http://en.wikipedia.org/wiki/Snow_Petrel

The text in this page is based on the copyrighted Wikipedia article shown in above URL. It is used under the GNU Free Documentation License. You may redistribute it, verbatim or modified, providing that you comply with the terms of the GFDL.